PyroClean: Denoising Pyrosequences from Protein-Coding Amplicons for the Recovery of Interspecific and Intraspecific Genetic Variation

	PyroClean: Denoising Pyrosequences from Protein-Coding Amplicons for the Recovery of Interspecific and Intraspecific Genetic Variation
	Ramirez-Gonzalez R 1; Yu DW 2,3; Bruce C 3; Heavens D 1; Caccamo M 1; Emerson BC*3; bemerson@ipna.csic.es
	2013
发表期刊	PLOS ONE
卷号	8 期号:3 页码:e57615
合作性质	其它
摘要	High-throughput parallel sequencing is a powerful tool for the quantification of microbial diversity through the amplification of nuclear ribosomal gene regions. Recent work has extended this approach to the quantification of diversity within otherwise difficult-to-study metazoan groups. However, nuclear ribosomal genes present both analytical challenges and practical limitations that are a consequence of the mutational properties of nuclear ribosomal genes. Here we exploit useful properties of protein-coding genes for cross-species amplification and denoising of 454 flowgrams. We first use experimental mixtures of species from the class Collembola to amplify and pyrosequence the 59 region of the COI barcode, and we implement a new algorithm called PyroClean for the denoising of Roche GS FLX pyrosequences. Using parameter values from the analysis of experimental mixtures, we then analyse two communities sampled from field sites on the island of Tenerife. Cross-species amplification success of target mitochondrial sequences in experimental species mixtures is high; however, there is little relationship between template DNA concentrations and pyrosequencing read abundance. Homopolymer error correction and filtering against a consensus reference sequence reduced the volume of unique sequences to approximately 5% of the original unique raw reads. Filtering of remaining non-target sequences attributed to PCR error, sequencing error, or numts further reduced unique sequence volume to 0.8% of the original raw reads. PyroClean reduces or eliminates the need for an additional, time-consuming step to cluster reads into Operational Taxonomic Units, which facilitates the detection of intraspecific DNA sequence variation. PyroCleaned sequence data from field sites in Tenerife demonstrate the utility of our approach for quantifying evolutionary diversity and its spatial structure. Comparison of our sequence data to public databases reveals that we are able to successfully recover both interspecific and intraspecific sequence diversity.
资助者	This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology.
收录类别	SCI
语种	英语
资助者	This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. ; This work was supported by a Research Fellowship to Brent Emerson from The Leverhulme Trust, project support from The Genome Analysis Centre (TGAC), the University of East Anglia, and the Consejo Superior de Investigaciones Cientı ´ficas. Doug Yu was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW- Z-1027, Y002731079), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology.
WOS记录号	WOS:000315897100027
引用统计	被引频次：12[WOS] [WOS记录] [WOS相关记录]
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/7418
专题	科研部门_生态学与环境保护中心（俞维理）遗传资源与进化国家重点实验室
通讯作者	bemerson@ipna.csic.es
作者单位	1.The Genome Analysis Centre, Norwich Research Park, Norwich, United Kingdom 2.Ecology, Conservation, and Environment Center, State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, People’s Republic of China 3.School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich, United Kingdom
推荐引用方式 GB/T 7714	Ramirez-Gonzalez R,Yu DW,Bruce C,et al. PyroClean: Denoising Pyrosequences from Protein-Coding Amplicons for the Recovery of Interspecific and Intraspecific Genetic Variation[J]. PLOS ONE,2013,8(3):e57615.
APA	Ramirez-Gonzalez R.,Yu DW.,Bruce C.,Heavens D.,Caccamo M.,...&bemerson@ipna.csic.es.(2013).PyroClean: Denoising Pyrosequences from Protein-Coding Amplicons for the Recovery of Interspecific and Intraspecific Genetic Variation.PLOS ONE,8(3),e57615.
MLA	Ramirez-Gonzalez R,et al."PyroClean: Denoising Pyrosequences from Protein-Coding Amplicons for the Recovery of Interspecific and Intraspecific Genetic Variation".PLOS ONE 8.3(2013):e57615.

条目包含的文件
文件名称/大小	文献类型	版本类型	开放类型	使用许可
2013050903.pdf（553KB）	期刊论文	出版稿	开放获取	CC BY-NC-SA	请求全文