Multiple Interkingdom Horizontal Gene Transfers in Pyrenophora and Closely Related Species and Their Contributions to Phytopathogenic Lifestyles

	Multiple Interkingdom Horizontal Gene Transfers in Pyrenophora and Closely Related Species and Their Contributions to Phytopathogenic Lifestyles
	Sun BF 1,2; Xiao JH 1; He SM 1; Liu L 1; Murphy RW 3,4; Huang DW*1,5; huangdw619@gmail.com
	2013
发表期刊	PLOS ONE
卷号	8 期号:3 页码:e60029
合作性质	其它
摘要	Many studies have reported horizontal gene transfer (HGT) events from eukaryotes, especially fungi. However, only a few investigations summarized multiple interkingdom HGTs involving important phytopathogenic species of Pyrenophora and few have investigated the genetic contributions of HGTs to fungi. We investigated HGT events in P. teres and P. tritici-repentis and discovered that both species harbored 14 HGT genes derived from bacteria and plants, including 12 HGT genes that occurred in both species. One gene coding a leucine-rich repeat protein was present in both species of Pyrenophora and it may have been transferred from a host plant. The transfer of genes from a host plant to pathogenic fungi has been reported rarely and we discovered the first evidence for this transfer in phytopathogenic Pyrenophora. Two HGTs in Pyrenophora underwent subsequent duplications. Some HGT genes had homologs in a few other fungi, indicating relatively ancient transfer events. Functional analyses indicated that half of the HGT genes encoded extracellular proteins and these may have facilitated the infection of plants by Pyrenophora via interference with plant defense-response and the degradation of plant cell walls. Some other HGT genes appeared to participate in carbohydrate metabolism. Together, these functions implied that HGTs may have led to highly efficient mechanisms of infection as well as the utilization of host carbohydrates. Evolutionary analyses indicated that HGT genes experienced amelioration, purifying selection, and accelerated evolution. These appeared to constitute adaptations to the background genome of the recipient. The discovery of multiple interkingdom HGTs in Pyrenophora, their significance to infection, and their adaptive evolution, provided valuable insights into the evolutionary significance of interkingdom HGTs from multiple donors
资助者	This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148.
收录类别	SCI
语种	英语
资助者	This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This project was supported by the National Natural Science Foundation of China (NSFC grant no. 31090253, 31210103912, 30900137), partially by Major Innovation Program of Chinese Academy of Sciences (KSCX2-EW-Z-2), a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences, National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0930004), and the Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148.
WOS记录号	WOS:000317263900053
引用统计	被引频次：30[WOS] [WOS记录] [WOS相关记录]
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/7426
专题	科研部门_分子进化与基因组多样性（张亚平）遗传资源与进化国家重点实验室
通讯作者	huangdw619@gmail.com
作者单位	1.Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China 2.University of the Chinese Academy of Sciences, Beijing, China 3.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China 4.Centre for Biodiversity and Conservation Biology, Royal Ontario Museum, Toronto, Canada 5.College of Plant Protection, Shandong Agricultural University, Tai’an, Shandong, China
推荐引用方式 GB/T 7714	Sun BF,Xiao JH,He SM,et al. Multiple Interkingdom Horizontal Gene Transfers in Pyrenophora and Closely Related Species and Their Contributions to Phytopathogenic Lifestyles[J]. PLOS ONE,2013,8(3):e60029.
APA	Sun BF.,Xiao JH.,He SM.,Liu L.,Murphy RW.,...&huangdw619@gmail.com.(2013).Multiple Interkingdom Horizontal Gene Transfers in Pyrenophora and Closely Related Species and Their Contributions to Phytopathogenic Lifestyles.PLOS ONE,8(3),e60029.
MLA	Sun BF,et al."Multiple Interkingdom Horizontal Gene Transfers in Pyrenophora and Closely Related Species and Their Contributions to Phytopathogenic Lifestyles".PLOS ONE 8.3(2013):e60029.

条目包含的文件
文件名称/大小	文献类型	版本类型	开放类型	使用许可
2013051604.pdf（1033KB）	期刊论文	出版稿	开放获取	CC BY-NC-SA	请求全文