KMS KUNMING INSTITUTE OF ZOOLOGY.CAS
| Spatial games and the maintenance of cooperation in an asymmetric Hawk-Dove game | |
| He JZ1,2; Zhao Y3; Cai HJ*3; Wang RW*2; hjcai@whu.edu.cn; wangrw@mail.kiz.ac.cn | |
| 2013 | |
| 发表期刊 | CHINESE SCIENCE BULLETIN
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| 卷号 | 58期号:18页码:2248-2254 特刊: SI |
| 合作性质 | 其它 |
| 摘要 | Classical theories explaining the evolution of cooperation often rely on the assumption that the involved players are symmetrically interacted. However, in reality almost all well-documented cooperation systems show that cooperative players are in fact asymmetrically interacted and that this dynamic may greatly affect the cooperative behavior of the involved players. Here, we developed several models based on the most well known spatial game of the Hawk-Dove game, while also considering the effects of asymmetric interaction. Such asymmetric games possess four kinds of strategies: cooperation or defection of strong player and cooperation or defection of weak player. Computer simulations showed that the probability of defection of the strong player decreases with decreasing the benefit to cost ratio, and that all kinds of strategy will be substituted by cooperation on behalf of the strong player if the benefit to cost ratio is sufficiently small. Moreover, weak players find it difficult to survive and the surviving weak players are mostly defectors, similar to the Boxed Pigs game. Interestingly, the patterns of kinds of strategies are chaotic or oscillate in some conditions with the related factors. |
| 关键词 | Asymmetric Interaction Cooperation Spatial Games Hawk-dove Game |
| 资助者 | This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). |
| 收录类别 | SCI |
| 语种 | 英语 |
| 资助者 | This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). ; This work was supported by the National Basic Research Program of China (2007CB411600), the National Natural Science Foundation of China (30670272, 30770500, 10961027, 31270433 and 10761010), the Yunnan Natural Science Foun- dation (2009CD104), the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF09-02), the West Light Foundation of the Chinese Academy of Sciences and Special Fund for the Excellent Youth of the Chinese Academy of Sciences (KSCX2-EW-Q-9). |
| WOS记录号 | WOS:000321629800023 |
| 引用统计 | |
| 文献类型 | 期刊论文 |
| 条目标识符 | http://ir.kiz.ac.cn/handle/152453/7599 |
| 专题 | 实验与理论生态学 |
| 通讯作者 | hjcai@whu.edu.cn; wangrw@mail.kiz.ac.cn |
| 作者单位 | 1.Statistics and Mathematics School, Yunnan University of Finance and Economics, Kunming 650221, China 2.Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, China 3.International School of Software, Wuhan University, Wuhan 430079, China |
| 推荐引用方式 GB/T 7714 | He JZ,Zhao Y,Cai HJ*,et al. Spatial games and the maintenance of cooperation in an asymmetric Hawk-Dove game[J]. CHINESE SCIENCE BULLETIN,2013,58(18):2248-2254 特刊: SI. |
| APA | He JZ,Zhao Y,Cai HJ*,Wang RW*,hjcai@whu.edu.cn,&wangrw@mail.kiz.ac.cn.(2013).Spatial games and the maintenance of cooperation in an asymmetric Hawk-Dove game.CHINESE SCIENCE BULLETIN,58(18),2248-2254 特刊: SI. |
| MLA | He JZ,et al."Spatial games and the maintenance of cooperation in an asymmetric Hawk-Dove game".CHINESE SCIENCE BULLETIN 58.18(2013):2248-2254 特刊: SI. |
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| 2013081509.pdf（1058KB） | 期刊论文 | 出版稿 | 开放获取 | CC BY-NC-SA | 请求全文 | |
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