Paternal origins of Chinese cattle

KIZ OpenIR > 科研部门 > 比较基因组学（宿兵）

	Paternal origins of Chinese cattle
	Li R 1; Zhang XM 2; Campana MG 3; Huang JP 1; Chang ZH 4; Qi XB2 ; Shi H 2; Su B2 ; Zhang RF 5; Lan XY 1; Chen H 1; Lei CZ*1; leichuzhao1118@126.com
	2013
发表期刊	ANIMAL GENETICS
卷号	44 期号:4 页码:446-449
合作性质	其它
摘要	To determine the genetic diversity and paternal origin of Chinese cattle, 302 males from 16 Chinese native cattle breeds as well as 30 Holstein males and four Burma males as controls were analysed using four Y-SNPs and two Y-STRs. In Chinese bulls, the taurine Y1 and Y2 haplogroups and indicine Y3 haplogroup were detected in seven, 172 and 123 individuals respectively, and these frequencies varied among the Chinese cattle breeds examined. Y2 dominates in northern China (91.4%), and Y3 dominates in southern China (90.8%). Central China is an admixture zone, although Y2 predominates overall (72.0%). The geographical distributions of the Y2 and Y3 haplogroup frequencies revealed a pattern of male indicine introgression from south to north China. The three Y haplogroups were further classified into one Y1 haplotype, five Y2 haplotypes and one Y3 haplotype in Chinese native bulls. Due to the interplay between taurine and indicine types, Chinese cattle represent an extensive reservoir of genetic diversity. The Y haplotype distribution of Chinese cattle exhibited a clear geographical structure, which is consistent with mtDNA, historical and geographical information.
关键词	Native Breeds Y-snp Y-str Haplotype Genetic Diversity
资助者	This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204).
收录类别	SCI
语种	英语
资助者	This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204). ; This work was supported by the Natural Science Founda- tion of China (31072001), the National ‘863’ Program of China (2008AA101010), the Program of National Beef Cattle Industrial Technology System (CARS-38), the Hubei Province Natural Science Foundation (2008CDB064) and the Natural Science Foundation of the Department of Education in Hubei Province (Q20082204).
WOS记录号	WOS:000329200100012
引用统计	被引频次：26[WOS] [WOS记录] [WOS相关记录]
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/7789
专题	科研部门_比较基因组学（宿兵）遗传资源与进化国家重点实验室
通讯作者	leichuzhao1118@126.com
作者单位	1.Shaanxi Key Laboratory of Molecular Biology for Agriculture, College of Animal Science and Technology, Northwest A&F University, Yangling, Shaanxi, 712100, China 2.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology and Kunming Primate Research Centre, Chinese Academy of Sciences, Kunming, Yunnan, 650223, China 3.Department of Human Evolutionary Biology, Harvard University, Cambridge, MA, 02138, USA 4.Weinan Vocational and Technical College, Weinan, Shaanxi, 714000, China 5.College of Life Science, National Experiment Teaching Center of Biology, Hubei Normal University, Huangshi, Hubei, 435002, China
推荐引用方式 GB/T 7714	Li R,Zhang XM,Campana MG,et al. Paternal origins of Chinese cattle[J]. ANIMAL GENETICS,2013,44(4):446-449.
APA	Li R.,Zhang XM.,Campana MG.,Huang JP.,Chang ZH.,...&leichuzhao1118@126.com.(2013).Paternal origins of Chinese cattle.ANIMAL GENETICS,44(4),446-449.
MLA	Li R,et al."Paternal origins of Chinese cattle".ANIMAL GENETICS 44.4(2013):446-449.

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