Mitochondrial phylogeography of the red-tailed knobby newt (Tylototriton kweichowensis)
Zhang MW*1; Yu GH2,3; Yang MX*1; Li Y1; Wei J1; Xu HL1; Yao YF1; Wang LJ1; Zhang XQ1; Rao DQ2; Yang JX2; mwzhangkiz@hotmail.com; cerana@yahoo.cn
2013
发表期刊BIOCHEMICAL SYSTEMATICS AND ECOLOGY
卷号51期号:X页码:195-202
合作性质其它
摘要The red-tail knobby newt (Tylototriton kweichowensis), endemic to Yunnan-Guizhou Plateau (China), is classed as vulnerable amphibian by IUCN. Recently, habitat destruction and degradation are major threats to this species. To assess the genetic diversity and population structure with this species, we determined the sequence of the mitochondrial DNA (mtDNA) control region and tRNA-Phe gene in 13 individuals from two sites in Yunnan, and reanalyzed 58 published samples from 12 sties in Yunnan and Guizhou. Nineteen variable sites were observed and 28 haplotypes were identified. Haplotype diversity was high (0.9364) and nucleotide diversity among all haplotypes was relatively low (0.00219). Bayesian tree and a median-joining network revealed no significant phylogeographic structure. Both the bell-shaped pairwise mismatch distribution and significant negative values of Fu's F-S suggested that red-tail knobby newt have experienced a recent population expansion. Base on the results of this study, several conservation recommendations on this vulnerable species are discussed.
关键词Conservation Tylototriton Kweichowensis Mitochondrial Dna Population Structure Vulnerable Species
资助者This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01).
收录类别SCI
语种英语
资助者This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01). ; This project was supported by Foundation of Education Department of Sichuan (13ZA0262), Shuang Zhi Ji Hua Foundation of Sichuan Agricultural University (01570709), Open projects of State Key Laboratory of Genetic Resources and Evolution, Kunming institute of Zoology, Chinese Academy of Sciences (GREKF11-01).
WOS记录号WOS:000329421000033
引用统计
文献类型期刊论文
条目标识符http://ir.kiz.ac.cn/handle/152453/7797
专题科研部门_系统进化与生物地理学(杨君兴)
遗传资源与进化国家重点实验室
科研部门_兽类生态与进化(蒋学龙)
通讯作者mwzhangkiz@hotmail.com; cerana@yahoo.cn
作者单位1.College of Animal Science and Technology, Sichuan Agricultural University, 625014 Ya’an, Sichuan, PR China
2.State Key laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, The Chinese Academy of Sciences, 32 Jiaochang Donglu, Kunming, Yunnan 650223, PR China
3.Graduate University of the Chinese Academy of Sciences, Beijing, PR China
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Zhang MW*,Yu GH,Yang MX*,et al. Mitochondrial phylogeography of the red-tailed knobby newt (Tylototriton kweichowensis)[J]. BIOCHEMICAL SYSTEMATICS AND ECOLOGY,2013,51(X):195-202.
APA Zhang MW*.,Yu GH.,Yang MX*.,Li Y.,Wei J.,...&cerana@yahoo.cn.(2013).Mitochondrial phylogeography of the red-tailed knobby newt (Tylototriton kweichowensis).BIOCHEMICAL SYSTEMATICS AND ECOLOGY,51(X),195-202.
MLA Zhang MW*,et al."Mitochondrial phylogeography of the red-tailed knobby newt (Tylototriton kweichowensis)".BIOCHEMICAL SYSTEMATICS AND ECOLOGY 51.X(2013):195-202.
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