| Isolation and characterization of thirteen microsatellite loci for the western black crested gibbon (Nomascus concolor) by high-throughput sequencing | |
| Hu NQ1,2; Joseph O3,4; Huang B1; He K1; Jiang XL*1; jiangxl@mail.kiz.ac.cn | |
| 2014 | |
| 发表期刊 | CONSERVATION GENETICS RESOURCES
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| 卷号 | 6期号:1页码:179-181 |
| 合作性质 | 其它 |
| 摘要 | The critically endangered western black crested gibbon (Nomascus concolor) lives only in isolated mountainous areas of China's Yunnan province, northern Vietnam, and northwestern Laos. A lack of genetic samples and polymorphic molecular loci has limited our ability to investigate the genetic diversity of this species. We collected fecal samples from 12 free-ranging individuals in the Wuliang Mountains of Yunnan and developed 13 polymorphic microsatellite loci using 1/8 plate of a Roche 454 GS-FLX sequencing run. The number of alleles ranged from three to eight per locus, and the observed and expected heterozygosities were 0.333-0.917 and 0.424-0.822, respectively. These loci will be useful tools not only for future studies of the social structure and population genetics of N. concolor, but also will facilitate an informed conservation management plan. |
| 关键词 | Western Black Crested Gibbon Nomascus Concolor Microsatellite High-throughput Sequencing |
| 资助者 | This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance |
| 收录类别 | SCI |
| 语种 | 英语 |
| 资助者 | This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance ; This study was supported by the National Nat- ural Science Foundation of China (31070349, 31170498), Yunnan Provincial (2011FB105), Chinese Academy of Science (KSCX2-EW- Z-4-1, GREKF11-04), the National Science Foundation (NSF OISE- 1015770, NSF BCS-1155904), The Leakey Foundation, The Ameri- can Philosophical Society Lewis and Clark Fund, Lambda Alpha, and Washington University in St. Louis. Thanks are given to The Key Lab of Animal Ecology and Conservation Biology, Institute of Zoology, The Chinese Academy of Sciences, for technical guidance |
| WOS记录号 | WOS:000331708300047 |
| 引用统计 | |
| 文献类型 | 期刊论文 |
| 条目标识符 | http://ir.kiz.ac.cn/handle/152453/7828 |
| 专题 | 科研部门_兽类生态与进化（蒋学龙） 遗传资源与进化国家重点实验室 |
| 通讯作者 | jiangxl@mail.kiz.ac.cn |
| 作者单位 | 1.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, 32 Jiaochang Donglu, Kunming, Yunnan 650223, People’s Republic of China 2.University of Chinese Academy of Sciences, Beijing, People’s Republic of China 3.State Key Laboratory of Genetic Resources and Evolution, Ecology, Conservation, and Environment Center (ECEC), Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, People’s Republic of China 4.Department of Anthropology, Washington University in St. Louis, St. Louis, MO, USA |
| 推荐引用方式 GB/T 7714 | Hu NQ,Joseph O,Huang B,et al. Isolation and characterization of thirteen microsatellite loci for the western black crested gibbon (Nomascus concolor) by high-throughput sequencing[J]. CONSERVATION GENETICS RESOURCES,2014,6(1):179-181. |
| APA | Hu NQ,Joseph O,Huang B,He K,Jiang XL*,&jiangxl@mail.kiz.ac.cn.(2014).Isolation and characterization of thirteen microsatellite loci for the western black crested gibbon (Nomascus concolor) by high-throughput sequencing.CONSERVATION GENETICS RESOURCES,6(1),179-181. |
| MLA | Hu NQ,et al."Isolation and characterization of thirteen microsatellite loci for the western black crested gibbon (Nomascus concolor) by high-throughput sequencing".CONSERVATION GENETICS RESOURCES 6.1(2014):179-181. |
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| 2014032802.pdf（161KB） | 期刊论文 | 出版稿 | 开放获取 | CC BY-NC-SA | 请求全文 | |
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