Avoidance of potentially harmful food cannot be socially transmitted between rats
Jing L1,2; Zhou QX*1; Xu L*1; lxu@vip.163.com; qixin_zhou@- 126.com
2014
发表期刊Zoological Research
卷号35期号:4页码:256−261
摘要The social transmission of food preferences (STFP) is a behavioural task of olfactory memory, in which an observer rat learns safe food odours from a demonstrator rat, and shows preference for this odour in a subsequent choice test. However, previous studies have failed to detect the transmission of information about food of potential danger and food aversion using STFP test. In this study, we tested how demonstrators' health affects the exchange of odour information and whether observers can learn danger information from an unhealthy demonstrator. As expected, the observer rat formed an odour preference after interacting with a demonstrator rat that had just eaten food containing a new odour, however, odour preference rather than aversion was also formed after interacting with a demonstrator rat injected with LiCl (used to induce gastric malaise). Furthermore, anaesthetized demonstrator rats and half-anaesthetized demonstrator rats, which showed obvious motor deficits suggesting an unhealthy state, also socially transmitted food preferences to observers. These results suggest that the social transmission of food preferences task is independent of a demonstrators' health, and that information about dangerous foods cannot be transmitted using this behavioural task.
关键词Diet Selection Rats Social Learning Social Transmission Of Food Preference
资助者This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775).
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语种英语
资助者This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775). ; This study was supported by the 973 Program of the Ministry of Science and Technology of China (2013CB835103); the Strategic Priority Research Program of Chinese Academy of Science (XDB02020200) and the National Science Foundation of China (31100775).
文献类型期刊论文
条目标识符http://ir.kiz.ac.cn/handle/152453/7937
专题科研部门_学习记忆的分子神经机制(徐林)
科研部门_动物模型与人类重大疾病机理重点实验室
通讯作者lxu@vip.163.com; qixin_zhou@- 126.com
作者单位1.Laboratory of Learning and Memory, Key Laboratory of Animal Models and Human Disease Mechanisms of the Chinese Academy of Sciences & Yunnan Province, Kunming Institute of Zoology, Kunming 650223, China
2.University of Chinese Academy of Sciences, Beijing 100049, China
推荐引用方式
GB/T 7714
Jing L,Zhou QX*,Xu L*,et al. Avoidance of potentially harmful food cannot be socially transmitted between rats[J]. Zoological Research,2014,35(4):256−261.
APA Jing L,Zhou QX*,Xu L*,lxu@vip.163.com,&qixin_zhou@- 126.com.(2014).Avoidance of potentially harmful food cannot be socially transmitted between rats.Zoological Research,35(4),256−261.
MLA Jing L,et al."Avoidance of potentially harmful food cannot be socially transmitted between rats".Zoological Research 35.4(2014):256−261.
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