Root parasitic plant Orobanche aegyptiaca and shoot parasitic plant Cuscuta australis obtained Brassicaceae-specific strictosidine synthase-like genes by horizontal gene transfer

	Root parasitic plant Orobanche aegyptiaca and shoot parasitic plant Cuscuta australis obtained Brassicaceae-specific strictosidine synthase-like genes by horizontal gene transfer
	Zhang DL 1,2; Qi JF 1; Yue JP 3,4; Huang JL 3,4; Sun T 1; Li SP 2; Wen JF5 ; Hettenhausen C 1; Wu JS 1; Wang L 1; Zhuang HF 1; Wu JQ1; Sun GL1; wujianqiang@mail.kib.ac.cn ; sung@mail.kib.ac.cn
	2014
发表期刊	BMC PLANT BIOLOGY
卷号	14 期号:X 页码:e19
摘要	Background: Besides gene duplication and de novo gene generation, horizontal gene transfer (HGT) is another important way of acquiring new genes. HGT may endow the recipients with novel phenotypic traits that are important for species evolution and adaption to new ecological niches. Parasitic systems expectedly allow the occurrence of HGT at relatively high frequencies due to their long-term physical contact. In plants, a number of HGT events have been reported between the organelles of parasites and the hosts, but HGT between host and parasite nuclear genomes has rarely been found. Results: A thorough transcriptome screening revealed that a strictosidine synthase-like (SSL) gene in the root parasitic plant Orobanche aegyptiaca and the shoot parasitic plant Cuscuta australis showed much higher sequence similarities with those in Brassicaceae than with those in their close relatives, suggesting independent gene horizontal transfer events from Brassicaceae to these parasites. These findings were strongly supported by phylogenetic analysis and their identical unique amino acid residues and deletions. Intriguingly, the nucleus-located SSL genes in Brassicaceae belonged to a new member of SSL gene family, which were originated from gene duplication. The presence of introns indicated that the transfer occurred directly by DNA integration in both parasites. Furthermore, positive selection was detected in the foreign SSL gene in O. aegyptiaca but not in C. australis. The expression of the foreign SSL genes in these two parasitic plants was detected in multiple development stages and tissues, and the foreign SSL gene was induced after wounding treatment in C. australis stems. These data imply that the foreign genes may still retain certain functions in the recipient species. Conclusions: Our study strongly supports that parasitic plants can gain novel nuclear genes from distantly related host species by HGT and the foreign genes may execute certain functions in the new hosts.
关键词	Strictosidine Synthase-like Genes Cuscuta Horizontal Gene Transfer New Genes Orobanche Parasitic Plants
资助者	This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group.
收录类别	SCI
语种	英语
资助者	This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group. ; This work is supported by a grant from the Natural Science Foundation of China (No. 31301037), a grant from the Natural Science Foundation of Yunnan Province of China (No. 2013FB068), a project sponsored by SRF for ROCS, SEM, a Startup grant from Kunming Institute of Botany, CAS, a grant from the Yunnan Recruitment Program of Experts in Sciences (No. 2012HA016), a grant from the Thousand Youth Talents Program, an Open Foundation of the State Key Laboratory of Genetics Resources and Evolution, CAS, and a grant for the Max Planck Partner Group.
WOS记录号	WOS:000330075900001
引用统计	被引频次：36[WOS] [WOS记录] [WOS相关记录]
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/8061
专题	科研部门_真核细胞进化基因组（文建凡）遗传资源与进化国家重点实验室
通讯作者	wujianqiang@mail.kib.ac.cn; sung@mail.kib.ac.cn
作者单位	1.Key Laboratory of Economic Plants and Biotechnology, Kunming Institute of Botany, Chinese Academy of Sciences, 132 Lanhei Road, Heilongtan, Kunming 650201, Yunnan, China 2.2 College of Life Science, Henan University, 85 Minglun Street, Kaifeng 475001, Henan, China 3.Key Laboratory of Biodiversity and Biogeography, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China 4.Department of Biology, East Carolina University, Greenville, NC 27858, USA 5.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, 32 Jiaochang East Road, Kunming 650223, Yunnan, China.
推荐引用方式 GB/T 7714	Zhang DL,Qi JF,Yue JP,et al. Root parasitic plant Orobanche aegyptiaca and shoot parasitic plant Cuscuta australis obtained Brassicaceae-specific strictosidine synthase-like genes by horizontal gene transfer[J]. BMC PLANT BIOLOGY,2014,14(X):e19.
APA	Zhang DL.,Qi JF.,Yue JP.,Huang JL.,Sun T.,...&sung@mail.kib.ac.cn.(2014).Root parasitic plant Orobanche aegyptiaca and shoot parasitic plant Cuscuta australis obtained Brassicaceae-specific strictosidine synthase-like genes by horizontal gene transfer.BMC PLANT BIOLOGY,14(X),e19.
MLA	Zhang DL,et al."Root parasitic plant Orobanche aegyptiaca and shoot parasitic plant Cuscuta australis obtained Brassicaceae-specific strictosidine synthase-like genes by horizontal gene transfer".BMC PLANT BIOLOGY 14.X(2014):e19.

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