| Niche Divergence Among Sex and Age Classes in Black-and-White Snub-nosed Monkeys (Rhinopithecus bieti) | |
| Wan Y1,2; Quan RC3; Ren GP1; Wang L1; Long YC4; Liu XH1; Zhu JG*1; zhu@mail.kiz.ac.cn | |
| 2013 | |
| 发表期刊 | INTERNATIONAL JOURNAL OF PRIMATOLOGY
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| 卷号 | 34期号:5页码:946-956 |
| 合作性质 | 其它 |
| 摘要 | A wide range of dimorphic species exhibit intraspecific differences in feeding ecology and habitat use; however, studies comparing resource partitioning between males and females and between age groups occur mainly at the community and species level. We here classified the main habitat of black-and-white snub-nosed monkeys (Rhinopithecus bieti) -fir tree crown- into six microhabitats; examined the lichen-load of the microhabitats; and investigated the microhabitat use of age-sex groups within one-male units (OMUs: an adult male, several females, and juveniles) of R. bieti using instantaneous scanning sampling methods to test whether there is resource niche partitioning in social units of this sexually dimorphic primate in a reserve in Yunnan, China. Our results showed that adult females and juveniles often foraged in the outer layer of the fir tree crown, whereas adult males often foraged in the inner layer, closer to the trunk where food abundance was highest. Adult females and juveniles, however, both shifted microhabitat-use patterns to the inner layer of the fir tree crown when the adult male was absent from the tree. We suggested that the preference for and benefit of the central layer of fir tree crown lies in food availability, but that adult male competitively excluded group members. Microhabitat segregation may serve to maximize the fitness of the group members by reducing intragroup competition, while also increasing interindividual distance, thereby reducing the possible effects of scramble competition. |
| 关键词 | Niche Partitioning Rhinopithecus Bieti Adult-juvenile Competition Habitat Male-female Competition |
| 资助者 | The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). |
| 收录类别 | SCI |
| 语种 | 英语 |
| 资助者 | The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). ; The study was supported by funds of the National Nature Science Foundation of China (31272327, 30770308), Committee for Research & Exploration, National Geographic Society of the USA (7962–05), The Nature Conservancy, and Kunming Institute of Zoology (Y106521131). |
| WOS记录号 | WOS:000325183200006 |
| 引用统计 | |
| 文献类型 | 期刊论文 |
| 条目标识符 | http://ir.kiz.ac.cn/handle/152453/8079 |
| 专题 | 科研部门_生态学与环境保护中心（俞维理） |
| 通讯作者 | zhu@mail.kiz.ac.cn |
| 作者单位 | 1.Ecology, Conservation, and Environment Center, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China 2.University of Chinese Academy of Sciences, Beijing 100101, China 3.Chinese Academy of Sciences Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Jinhong, Yunnan 666303, China 4.The Nature Conservancy, China Program, Kunming, Yunnan 650021, China |
| 推荐引用方式 GB/T 7714 | Wan Y,Quan RC,Ren GP,et al. Niche Divergence Among Sex and Age Classes in Black-and-White Snub-nosed Monkeys (Rhinopithecus bieti)[J]. INTERNATIONAL JOURNAL OF PRIMATOLOGY,2013,34(5):946-956. |
| APA | Wan Y.,Quan RC.,Ren GP.,Wang L.,Long YC.,...&zhu@mail.kiz.ac.cn.(2013).Niche Divergence Among Sex and Age Classes in Black-and-White Snub-nosed Monkeys (Rhinopithecus bieti).INTERNATIONAL JOURNAL OF PRIMATOLOGY,34(5),946-956. |
| MLA | Wan Y,et al."Niche Divergence Among Sex and Age Classes in Black-and-White Snub-nosed Monkeys (Rhinopithecus bieti)".INTERNATIONAL JOURNAL OF PRIMATOLOGY 34.5(2013):946-956. |
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