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KIZ OpenIR  > 科研部门  > 比较基因组学（宿兵）
Evolutionary Origin and Human-Specific Expansion of a Cancer/Testis Antigen Gene Family
Zhang Q*1; Su B*2
2014
发表期刊MOLECULAR BIOLOGY AND EVOLUTION
卷号31期号:9页码:2365-2375
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摘要

Cancer/testis (CT) antigens are encoded by germline genes and are aberrantly expressed in a number of human cancers. Interestingly, CT antigens are frequently involved in gene families that are highly expressed in germ cells. Here, we presented an evolutionary analysis of the CTAGE (cutaneous T-cell-lymphoma-associated antigen) gene family to delineate its molecular history and functional significance during primate evolution. Comparisons among human, chimpanzee, gorilla, orangutan, macaque, marmoset, and other mammals show a rapid and primate specific expansion of CTAGE family, which starts with an ancestral retroposition in the haplorhini ancestor. Subsequent DNA-based duplications lead to the prosperity of single-exon CTAGE copies in catarrhines, especially in humans. Positive selection was identified on the single-exon copies in comparison with functional constraint on the multiexon copies. Further sequence analysis suggests that the newly derived CTAGE genes may obtain regulatory elements from long terminal repeats. Our result indicates the dynamic evolution of primate genomes, and the recent expansion of this CT antigen family in humans may confer advantageous phenotypic traits during early human evolution.

关键词Ctage Gene Family Cancer/testis Antigen Positive Selection Gene Duplication Retroposition Human-specific Expansion
资助者The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002).
收录类别SCI
语种英语
资助者The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002). ; The authors appreciate helpful comments from anonymous reviewers. This work was supported by the Department of Human Evolutionary Biology, Harvard University, by grants fromtheNational973projectofChina(2011CBA00401),and theNationalNaturalScienceFoundationofChina(31130051, 31301028, and 31321002).
WOS记录号WOS:000343401100010
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文献类型期刊论文
条目标识符http://ir.kiz.ac.cn/handle/152453/8176
专题科研部门_比较基因组学（宿兵）
遗传资源与进化国家重点实验室
作者单位1.Department of Human Evolutionary Biology, Graduate School of Art and Science, Harvard University
2.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China
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GB/T 7714		 Zhang Q*,Su B*. Evolutionary Origin and Human-Specific Expansion of a Cancer/Testis Antigen Gene Family[J]. MOLECULAR BIOLOGY AND EVOLUTION,2014,31(9):2365-2375.
APA Zhang Q*,&Su B*.(2014).Evolutionary Origin and Human-Specific Expansion of a Cancer/Testis Antigen Gene Family.MOLECULAR BIOLOGY AND EVOLUTION,31(9),2365-2375.
MLA Zhang Q*,et al."Evolutionary Origin and Human-Specific Expansion of a Cancer/Testis Antigen Gene Family".MOLECULAR BIOLOGY AND EVOLUTION 31.9(2014):2365-2375.
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