| Multilocus species delimitation in the Crotalus triseriatus species group (Serpentes: Viperidae: Crotalinae), with the description of two new species | |
| Bryson RW*1; Linkem CW1; Dorcas ME2; Lathrop A3; Jones JM4; Alvarado-Diaz J5; Grusnwald CI6; Murphy RW3,7; brysonjr@uw.edu | |
| 2014 | |
| 发表期刊 | ZOOTAXA
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| 卷号 | 3826期号:3页码:475-496 |
| 合作性质 | 其它 |
| 摘要 | Members of the Crotalus triseriatus species group of montane rattlesnakes are widely distributed across the highlands of Mexico and southwestern USA. Although five species are currently recognized within the group, species limits remain to be tested. Genetic studies suggest that species may be paraphyletic and that at least one cryptic species may be present. We generate 3,346 base pairs of DNA sequence data from seven nuclear loci to test competing models of species delimitation in the C. triseriatus group using Bayes factor delimitation. We also examine museum specimens from the Trans-Mexican Volcanic Belt for evidence of cryptic species. We find strong support for a nine-species model and genetic and morphological evidence for recognizing two new species within the group, which we formally describe here. Our results suggest that the current taxonomy of the C. triseriatus species group does not reflect evolutionary history. We suggest several conservative taxonomic changes to the group, but future studies are needed to better clarify relationships among species and examine genetic patterns and structure within wide-ranging lineages. |
| 关键词 | Bayes Factor Delimitation Cloud Forest Mexico Trans-mexican Volcanic Belt |
| 资助者 | This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. |
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| 资助者 | This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. ; This paper is based in part upon work supported by the National Science Foundation grants DEB- 1257785 and DEB- 1258205 and by Natural Sciences and Engineering Research Council (Canada) Discovery Grant 3148. |
| WOS记录号 | WOS:000338113400003 |
| 引用统计 | |
| 文献类型 | 期刊论文 |
| 条目标识符 | http://ir.kiz.ac.cn/handle/152453/8182 |
| 专题 | 科研部门_分子进化与基因组多样性（张亚平） 遗传资源与进化国家重点实验室 |
| 通讯作者 | brysonjr@uw.edu |
| 作者单位 | 1.Department of Biology & Burke Museum of Natural History and Culture, University of Washington, Box 351800, Seattle, WA 98195- 1800, USA. E-mail: brysonjr@uw.edu 2.Department of Biology, Davidson College, Davidson, NC 28035-7118, USA. E-mail: midorcas@davidson.edu 3.Centre for Biodiversity and Conservation Biology, Royal Ontario Museum, Toronto, Ontario, Canada M5S 2C6. E-mail: amyl@rom.on.ca; bob.murphy@utoronto.ca 4.16310 Avenida Florencia, Poway, California 92064, USA. E-mail: jasonjones@crotalus.com 5.Instituto de Investigaciones sobre los Recursos Naturales, Universidad Michoacana de San Nicolás de Hidalgo, Av. San Juanito Itzic- uaro s/n, Col. Nueva Esperanza, C.P. 58337, Morelia, Michoacán, México. 6.Carretera Chapala-Jocotepec Oriente #57-1, Col. Centro, C.P. 45920, Ajijic, Jalisco, México. E-mail: cgruenwald@switaki.com 7.State Key Laboratory of Genetic Resources and Evolution, and Yunnan Laboratory of Molecular Biology of Domestic Animals, Kun- ming Institute of Zoology, Chinese Academy of Sciences, 32 Jiaochang Donglu, Kunming 250223, Yunnan, China |
| 推荐引用方式 GB/T 7714 | Bryson RW*,Linkem CW,Dorcas ME,et al. Multilocus species delimitation in the Crotalus triseriatus species group (Serpentes: Viperidae: Crotalinae), with the description of two new species[J]. ZOOTAXA,2014,3826(3):475-496. |
| APA | Bryson RW*.,Linkem CW.,Dorcas ME.,Lathrop A.,Jones JM.,...&brysonjr@uw.edu.(2014).Multilocus species delimitation in the Crotalus triseriatus species group (Serpentes: Viperidae: Crotalinae), with the description of two new species.ZOOTAXA,3826(3),475-496. |
| MLA | Bryson RW*,et al."Multilocus species delimitation in the Crotalus triseriatus species group (Serpentes: Viperidae: Crotalinae), with the description of two new species".ZOOTAXA 3826.3(2014):475-496. |
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| 文件名称/大小 | 文献类型 | 版本类型 | 开放类型 | 使用许可 | ||
| 2014112401.pdf（6525KB） | 期刊论文 | 出版稿 | 开放获取 | CC BY-NC-SA | 请求全文 | |
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