学习记忆的分子神经机制（徐林）知识库

QQ客服 官方微博 反馈留言

	Fluoxetine treatment reverses the intergenerational impact of maternal separation on fear and anxiety behaviors
	Xiong GJ1,2; Yang Y3; Cao J1; Mao RR*1; Xu L*1; talktomaomao@163.com
	2015
发表期刊	NEUROPHARMACOLOGY
卷号	92期号:X页码:1-7
合作性质	其它
摘要	Early life stress increase's risks of fear and anxiety related disorders in adulthood, which may be alleviated by fluoxetine treatment. However, the intergenerational impacts of maternal separation (MS) on fear and anxiety behaviors from father to their offspring are little known. And the potential effects of fluoxetine treatment on the intergenerational transmission have not been well tested. Here, we investigated whether fluoxetine can reverse the intergenerational effects of MS on fear and anxiety behaviors. The first generation (F1) male rats were exposed to MS 3 h daily from postnatal day 2-14 and then treated with fluoxetine for four weeks during adulthood before fear conditioning. We found that maternal separation significantly impaired contextual fear extinction in F1 adult male rats but not in their second generation (F2). Although no obvious effects of MS on anxiety were observed in F1 male rats, the F2 offspring displayed a phenotype of low anxiety-like behaviors despite they were reared in normal condition. Fluoxetine treatment in F1 males not only reversed the impairment of fear extinction in F1 males but also the low anxiety-like behaviors in their F2 offspring. These findings highlight the intergenerational impacts of early life stress on fear and anxiety behaviors, and provide a new sight of the intergenerational effect of fluoxetine therapy for early life stress related mental problems.
关键词	Fluoxetine Maternal Separation Fear Extinction Offspring Intergenerational Anxiety
资助者	This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048).
收录类别	SCI
语种	英语
资助者	This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048). ; This work was supported by National Basic Research Program of China (2013CB835103), and Strategic Priority Research Program of the Chinese Academy of Sciences (XDB02020002), NSFC-CIHR Joint Grant (81161120536), National Natural Science Foundation of China (31100786, 81171294, U1032605 and U1132602), Science and Technology Program of Yunnan Province (2013GA003 and 2013FA048).
WOS记录号	WOS:000351793700001
引用统计	被引频次：9[WOS]   [WOS记录]     [WOS相关记录]
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/8309
专题	科研部门_学习记忆的分子神经机制（徐林） 科研部门_动物模型与人类重大疾病机理重点实验室
通讯作者	talktomaomao@163.com
作者单位	1.Key Laboratory of Animal Models and Human Disease Mechanisms, and KIZ/CUHK Joint Laboratory of Bioresources and Molecular Research in Common Disease, and Laboratory of Learning and Memory, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, Yunnan, China 2.University of Chinese Academy of Sciences, Beijing 100049, China 3.Department of Physiology, Kunming Medical University, Kunming 650500, Yunnan, China
推荐引用方式 GB/T 7714	Xiong GJ,Yang Y,Cao J,et al. Fluoxetine treatment reverses the intergenerational impact of maternal separation on fear and anxiety behaviors[J]. NEUROPHARMACOLOGY,2015,92(X):1-7.
APA	Xiong GJ,Yang Y,Cao J,Mao RR*,Xu L*,&talktomaomao@163.com.(2015).Fluoxetine treatment reverses the intergenerational impact of maternal separation on fear and anxiety behaviors.NEUROPHARMACOLOGY,92(X),1-7.
MLA	Xiong GJ,et al."Fluoxetine treatment reverses the intergenerational impact of maternal separation on fear and anxiety behaviors".NEUROPHARMACOLOGY 92.X(2015):1-7.

条目包含的文件
文件名称/大小	文献类型	版本类型	开放类型	使用许可
2015042402.pdf（776KB）	期刊论文	出版稿	开放获取	CC BY-NC-SA	请求全文

个性服务

推荐该条目

保存到收藏夹

查看访问统计

导出为Endnote文件

谷歌学术

谷歌学术中相似的文章

[Xiong GJ]的文章

[Yang Y]的文章

[Cao J]的文章

百度学术

百度学术中相似的文章

[Xiong GJ]的文章

[Yang Y]的文章

[Cao J]的文章

必应学术

必应学术中相似的文章

[Xiong GJ]的文章

[Yang Y]的文章

[Cao J]的文章

相关权益政策

暂无数据

收藏/分享

除非特别说明，本系统中所有内容都受版权保护，并保留所有权利。